Botanists from Real Jardín Botánico of Madrid, Missouri Botanical Garden and University of Murcia carried out an expedition to Ecuador in July and August of 2006. The briological team was composed by María J. Cano, M. Teresa Gallego, Juan Guerra and Juan A. Jiménez (University of Murcia), and it was funded by Spanish Ministry of Education and Science through the projects “Taxonomic revision of the genus Hennediella, Syntrichia and Tortula (Pottiaceae, Bryophyta) in South America” (CGL2004-00788) and “Flora Briofítica Ibérica” (REN2003-00766). This exploration was carried out thanks to the cooperation of National University of Loja. We wish to acknowledge the assistance of Danilo Minga (University of Azuay, Cuenca), Susana León (Catholic University of Quito) and Luis A. Gutiérrez (National University of Loja). About 600 specimens of bryophytes were collected, keeping primarily at MUB with duplicates at LOJA, and MO.



Pichincha: reserva de Yanacocha, 00º06'38''S 78º34'34''W, 3360 m.
Pichincha: carretera hacia la reserva de Yanacocha, 00º06'16''S 78º31'52''W, 3220 m.


Pichincha: reserva geobotánica Pululahua, 00º01'31''S 78º29'00''W, 2795 m.
Pichincha: San Antonio de Pichincha, 00º01'42''S 78º26'10''W, 2480 m.
Pichincha: parque Jerusalén, 00º00'02''N 78º21'21''W, 2340 m.
Pichincha: pr. parque Jerusalén, 00º00'44''S 78º20'34''W, 2300 m.


Pichincha: Reserva Cayanbe-Coca, vía a Baeza, 00º00'44''S 78º20'34''W, 4200 m.
Pichincha: Peñas Blancas, 00º18'10''S 78º14'27''W, 3720 m.


Chimborazo: pr. Chunchi, en la Panamericana sur, 02º15'58''S 78º53'28''W, 2440 m.




Azuay: Santa Isabel, desvío a Pucará,03º19'47''S 78º22'52''W, 1120 m.
Azuay: río San Francisco, carretera Pasaje-Cuenca, 03º18'51''S 78º29'02''W, 825 m.


Loja: sur de San Lucas, 03º47'51''S 79º15'38''W, 2490 m.
Loja: Loja, plaza de San Sebastian, 04º02'S 79º12''W, 2100 m.


Loja: Parque Nacional Podocarpus, Cajanuma, 04º06'58''S 79º10'14''W, 3375 m.


Loja: Parque Nacional Podocarpus, Cerro Toledo, pr. Yangana, 04º22'15''S 79º09'28''W, 2190 m.
Loja: Parque Nacional Podocarpus, Cerro Toledo, 04º22'29''S 79º06'40'''W, 3110 m.
Loja: Parque Nacional Podocarpus, Cerro Toledo, pr. Yangana, 04º23'02''S 79º07'00''W, 3067 m.


Loja: pr. Velacruz, 03º59'55''S 79º35'25''W, 1920 m.
Loja: pr. Catacocha, 04º02'06''S 79º41'10''W, 1070 m.
Loja: pr. Casanga, 04º00'23''S 79º45'52''W, 1200 m.
Loja: pr. Catacocha, 04º01'54''S 79º41'28''W, 925 m.
Loja: pr. Velacruz, 03º58'28''S 79º30'17''W, 1250 m.


Cuenca: Baños, Hostería Durán, 02º55'12''S 79º03'42''W, 2680 m.
Chimborazo: pr. Palmira, 02º07'49''S 79º44'29''W, 3500 m.
Chimborazo: pr. Palmira, 02º07'49''S 79º44'29''W, 3500 m.


Chimborazo: ladera sur del volcán Chimborazo, 01º31'15''S 78º50'48''W, 4010 m.
Chimborazo: ladera sur del volcán Chimborazo, 01º29'45''S 78º51'50''W, 4370 m.
Chimborazo: ladera sur del volcán Chimborazo, 01º29'00''S 78º51'57''W, 4580 m.
Chimborazo: ladera sur del volcán Chimborazo, sobre el segundo refugio, 01º28'22''S 78º50'19''W, 5040 m.


Chimborazo: pr. San Juan, 01º37'02''S 78º48'41''W, 3430 m.
Chimborazo: páramo de Tililac, pr. San Juan, 01º38'52''S 78º50'51''W, 4000 m.
Chimborazo: carretera Riobamba-Guaranda, 01º31'50''S 78º49'46''W, 3800 m.


Cotopaxi: Hacienda La Ciénega, 00º46'24''S 78º36'51''W, 2945 m.
Cotopaxi: Parque Nacional Cotopaxi, volcán Cotopaxi, camino al refugio José F. Rivas,00º39'00''S 78º26'11''W, 4415 m.
Cotopaxi: Parque Nacional Cotopaxi, pr. Ingapirca,00º35'32''S 78º26'21''W, 3750 m.



  • Jiménez J.A., Ros R.M., Cano M.J. & Guerra J. 2005. A new evaluation of the genus Trichostomopsis (Pottiaceae, Bryophyta). Botanical Journal of the Linnean Society 147: 117–127.
  • Gallego M.T. & Cano M.J. 2007. New reports of Syntrichia Brid. (Pottiaceae, Bryophyta) taxa from South America. Nova Hedwigia 85: 457–461.
  • Jiménez J.A. 2007. Bryoerythrophyllum berthoanus comb. nov. and B. sharpii (Bryophyta, Pottiaceae) in South America. Journal of Bryology 29:125–130.
  • Cano M.J. 2008. Taxonomic revision of Hennediella Paris (Pottiaceae, Bryophyta). Bryophytorum Bibliotheca. 64. J. Cramer, Berlin.
  • Cano M.J. & Gallego M.T., Jiménez J.A. & Guerra J. 2008. Aloina obliquifolia (Pottiaceae, Bryophyta) new to South America, and new reports of Aloina in the Neotropics. Cryptogamie, Bryologie 29: 75–81.
  • Cano M.J., Jiménez J.A., Churchill S.P. & Guerra J. 2008. New records of Pottiaceae (Bryophyta) for South America. Cryptogamie, Bryologie 29: 183–191.
  • Jiménez J.A. & Cano M.J. 2008. Taxonomic assessment of Didymodon pruinosus (Mitt.) R.H. Zander (Bryophyta, Pottiaceae) a poorly known South American taxon. Nova Hedwigia 87: 145–152.
  • Jiménez J.A., Cano M.J., Guerra J., Churchill S.P. & Suárez G.M. 2009. Pseudocrossidium linearifolium (Potttiaceae) comb. et stat. nov. The Bryologist 112: 188–193.
  • Cano M.J., Jiménez J.A. & Jiménez J.F. 2010. A systematic revision of the genus Erythrophyllopsis (Pottiaceae, Bryophyta). Systematic Botany 35: 683–694.
  • Jiménez J.A., Cano M.J. Gallego M.T. & Guerra J. 2012. Didymodon lainzii (Pottiaceae) a new species from the tropical seasonally dry forest of Ecuador and Peru. The Bryologist 115: 406–411.
  • Cano M.J. & Jiménez J.A. 2013. A taxonomic revision of the tribe Pleuroweisieae (Pottiaceae, Bryophyta) in South America. Phytotaxa 143: 1–42. 
  • Jiménez J.A. & Cano M.J. 2014. A new species of Didymodon (Bryophyta, Pottiaceae), from northern South America. Nova Hedwigia 99: 467–473. 
  • Alonso M., Jiménez J. A. & Cano M. J. 2019. Taxonomic revision of Chionoloma (Pottiaceae, Bryophyta). Annals of the Missouri Botanical Garden 104: 563–632.
  • Cano M.J., Jiménez J.A., Gallego M.T. & Guerra J. 2021. A molecular approach to the phylogeny of the moss genus Pseudocrossidium (Pottiaceae, Bryopsida). Journal of Systematics and Evolution, jse.12801.
  • Jiménez J.A., Cano M.J. & Guerra J. 2022 [2021]. A multilocus phylogeny of the moss genus Didymodon and allied genera (Pottiaceae): Generic delimitations and their implications for systematics. Journal of Systematics and Evolution 60(2): 281–304.
  • M.T. Gallego, M.J. Cano , J.A. Jiménez & J. Guerra. 2022. Circumscription and Phylogenetic Position of Two Propagulose Species of Syntrichia (Pottiaceae, Bryophyta) Reveals Minor Realignments within the Tribe Syntricheae. Plants 11(5): 626.